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Moreover, p53 can increase the expression of spermine/spermine N1 acetyltransferase 1 (SAT1) and glutaminase 2 (GLS2), making cells more sensitive to ferroptosis, or promote ferroptosis by transcriptionally upregulating mitochondrial GLS2 [29]

This proposed hierarchical framework requires validation through systematic metabolomic and functional studies to establish the relative contributions of different metabolites, thereby informing targeted therapeutic strategies (Figure 1)

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