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Some proteins have layered stacks of -sheets, which impart structural integrity and may open to form a cavity (a beta barrel)

Similar nucleotide replacements and deletions are also observed in spontaneous overproducing isolates, including the vaginal isolate Limosilactobacillus reuteri AMBV339, that can reach high riboflavin levels of approximately 18.36 g/mL in laboratory conditions and food matrices 36

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Compared with fulllength TB500, the fragment shares the same peptide sequence (LKKTETQ) but lacks an aldehyde group on the leucine residue, a difference that reduces its molecular weight (889.0 vs 846.97 g/mol) and is proposed to increase stability, prolong halflife, and reduce offtarget interactions